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In addition to starch and protein, the starchy endosperm is a repository for nucleic acids and micronutrients.
From this it can be concluded that it takes 5 days for all the starchy endosperm cells to lose plasma membrane integrity.
Since they do not surround starchy tissue which requires hydrolysis during germination, it is unlikely that this aleurone produces large quantities of -amylases.
The secretory apparatus of the aleurone then proliferates, supporting increased hydrolase synthesis and secretion to degrade the starchy endosperm.
The amylases that degrade the starchy endosperm after germination are largely secreted by the aleurone layer after imbibition.
However, we will come back to the problem of small starchy seeds later.
Thus, there is no clear ultrastructural characterization of how starchy endosperm cells die during development.
Three classes of aleurone cells are distinct from the typical aleurone surrounding most of the starchy endosperm.
No expression is found in the parts of the seed not surviving desiccation, the maternal tissue and the starchy endosperm.
Surrounding the starchy endosperm is a highly specialized layer of endosperm tissue that remains cellular and viable at grain maturity.
Other components of the starchy endosperm may be further regulators of aleurone function.
The latter type probably characterizes dead cells because of the similarity in appearance to mature starchy endosperm cells (micrograph not shown).
Thus, in the case of premature slow drying, when differentiation into starchy endosperm cells is arrested, the meristematic cells acquire desiccation tolerance.
The endo-b-mannanase in the starchy endosperm was mainly from the aleurone layer.
The storage polymers in the starchy endosperm must then be mobilized and transported to the embryo to fuel its subsequent heterotrophic growth.
