The relative fitnesses are also scaled by the arbitrary assumption of a generation time l 15 days made when fitting the discrete model.
If the wild-type homozygote is viable and fertile, then it is impossible to infer the four fitnesses from the three chromosome frequencies alone.
346 sume the one-locus model of fitnesses given above.
However, locally stable polymorphisms are possible even if all double homozygotes have higher geometric mean fitnesses than all other genotypes.
If so, then mean fitnesses do not climb peaks in the adaptive topography.
The fitnesses of the heterozygotes and homozygotes for the disfavoured allele are 1khs and 1ks, respectively.
Offspring phenotypes and fitnesses are assigned as above.
Also, relative fitnesses might change through time, as a result of direct or indirect frequency-dependence.
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