They have four toes on their hind feet with claws that look like small hoofs; these are wider than those found on the forefeet.
The forefeet are large with powerful digging claws, while the hindfeet are stout, with flat soles.
The fifth toe on the forefeet is vestigial, consisting only of a tiny claw located on a tubercle at the side of the foot.
These monkeys have a unique feature in that their thumb is severely reduced on their forefeet, while the hindfeet retain five digits.
The forefeet possess four clawed digits, the third digit bearing the largest claw, while the hind feet have five digits.
Scent glands, which secrete a clear oil, are present in the forefeet and are larger in males than females.
Unlike the first, this trackway preserved traces of the forefeet.
The forefeet are heavily adapted for digging, with powerful curved claws.
