These are generally minor physical anomalies ranging from webbed toes to altered craniofacial morphology.
Fingers could never be discernible at the distance involved; this is consistent with the webbed fingers of the description.
We propose, instead, that the pes print morphology indicates that the track-makers may have had webbed feet.
Nor are there any seabirds with webbed feet.
The second and third, and fourth and fifth toes were webbed bilaterally, and the nails were small.
It is easy to build such webbed models.
More generally, this is true of infinitely many webbed models.
Graph models form the simplest class of webbed models.