Caryonides are lineages of paramecia that derive their macronuclei (somatic nuclei) from single macronuclear primordia (anlagen).
He wrote on the memory image, analysis of complex sensations, learning in paramecia, mental disorders, and anthropological psychology.
When macronuclei of clonally young "paramecia" were injected into "paramecia" of standard clonal age, the lifespan (clonal fissions) of the recipient was prolonged.