The manus impressions are tridactyl, strongly asymmetric, and rotated outward at almost a right angle to the long axis of the track way.
The manus impression is tridactyl, strongly asymmetric, and rotated outward at almost a right angle to the long axis of the trackway.
The manus print is about 330 mm in length, and 110 mm in width, consequently, the manus and pes impressions are of similar length.
Details of the prints suggest that the pes and possibly the manus bore webs of skin between the digits.
The digitigrade tridactyl manus impressions exhibit features of a typical pterosaur hand print.
It can be divided into the upper arm "brachium", the forearm "antebrachium", and the hand "manus".
The manus (hand) was composed merely of the five metacarpalia, which were orientated vertically and arranged in a semicircle.
This was probably a hang-over from a time when "dominium" (ownership), "manus" and "potestas" were indistinct and not formally separate.
