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Myriapods are arthropods with one pair of antennae, two or three pairs of mouthparts and many pairs of legs.
His paper studies myriapod segmentation.
It also plays host to a number of geological 'oddities' including fulgurites and giant myriapod trails.
It is the first myriapod, and the oldest known creature to have lived on land.
Previous to this he had suggested that a large crustacean, annelid worm or myriapod (such as a millipede) could have made them.
The myriapod head is very similar to that of the insects.
Due to the economic crisis during the late 1920s he had to retire from any bureaucratic activity and focussed exclusively on his myriapod studies.
Myriapod mouthparts are similar to those of chewing insects, although there is some variation between the myriapod classes.
